SAN FRANCISCO (AP) – Floyd, the oldest and tallest giraffe at the San Francisco Zoo, died Friday.

Floyd the giraffe. (photo credit: Marianne Hale)

The 18-year-old male reticulated giraffe had joint degeneration from aging and had suffered a “significant decline in health,” a zoo statement said.

“His imposing size yet serene demeanor, and independent spirit will be missed,” the zoo said.

Floyd was 16 feet (4.88 meters) tall and weighed more than 2,425 pounds (1,100 kilograms).

He was born in 2002 at the Albuquerque Zoo and transferred to San Francisco in 2004.

Over his lifetime, Floyd produced 11 offspring and had 24 grandchildren and five great-grandchildren who can be found all over the United States, the zoo said.

“Floyd was truly one of our most beloved animals and a favorite among our guests because he was so recognizable and engaging,” said Tanya M. Peterson, chief executive officer and executive director of the San Francisco Zoological Society, said in the statement. “We are heartbroken but know that he has touched so many during his life, and contributed greatly to the population of reticulated giraffes in North America.”

The reticulated giraffe is considered a vulnerable species by the International Union for Conservation of Nature. Poaching has reduced its native African population by half, according to the zoo.

Copyright 2020 The Associated Press.